Invertebrate survey at Gunpowder Park, the former Royal Ordnance Site in the Lee Valley Proposed management for invertebrates Invertebrate species may occur in a number of different habitats, they may have a range of differing ecological requirements. Many species are likely to have complex life cycles, with different specific requirements at different stages of their life cycle, but these requirements arc only known for rather few species. This indicates the importance, especially to the rare and scarcer species, of a continuity of habitat mosaic within an area. The requirement of many invertebrate species for a suitable foraging area where nectar and pollen may be gathered is paramount to their survival. Whilst many species may forage on a range of flower species, others are highly plant-specific. In addition to forage areas, predatory species require suitable hunting areas of open flower-rich vegetation, grassland or scrub that provide the specific prey species and it seems probable that fairly extensive areas are needed by some species. There is a parallel need for suitable larval development or nesting habitat and, in the case of parasitic species, there is a requirement for the presence of the host Larval development may take place on or in roots, stems, leaves, buds and seeds of plants or nesting may take place in the ground. For stem nesting species it is critical that these resources remain in situ from one generation to the next for the survival of these species and their parasites. Open bare areas are an important and often undervalued resource. Bare soil offers a number of benefits. In particular it warms up rapidly in sunshine, it is used by burrowing and ground nesting species and for egg laying and it provides a clear visual field for predators (Key 2000). Many species are associated with warm habitats, short dry grassland and disturbed ground. Sparsely vegetated and bare ground areas arc valuable components of these habitats, and sandy substrates are especially important to many aculeate Hymenoptera. South facing slopes and banks arc important features providing conditions favoured by thermophilic species at the edge of their range in Britain. The aspect and slope help promote summer drought and the retention of open areas. Even quite small areas can be important to ground nesting species. The diversity of flower-rich vegetation supports a variety of invertebrates that have feeding relationships with the different plant species. The variety of flowers is an important forage source for species of aculeate Hymenoptera and for invertebrates in general. The importance lies not only with the range of flower species present but the continuity throughout the season. This is crucial to bumblebees, which depend on a continuity of suitable forage sources throughout the spring and summer for nest success, but also in providing specific pollen and nectar resources for different species of aculeate during the year - many solitary bee species have short seasons where they are dependent on specific forage species. Many grassland invertebrates of conservation interest develop inside, feed on or over winter in seeds and fruit heads and are therefore dependent on a continuity of these resources. The invertebrate fauna is largely dependent on a low level of regular management combined with occasional disturbance to enable the survival of species that develop inside leaves, stems, flower heads and seeds and which over winter inside or at the bases of dead herbaceous stems and in ground litter. For stem nesting species it is critical that these resources remaining in situ from one generation to the next for the survival of these species and their cleptoparasites. 108 Essex Naturalist (New Series) 21 (2004)